Formosan Subterranean Termite1
B. J. Cabrera, N.-Y. Su, R. H. Scheffrahn, F. M. Oi,
and P. G. Koehler
2
Introduction
The Formosan subterranean termite, Coptotermes
formosanus Shiraki, is one of the most destructive termite
species in the world today. In the United States it causes
tremendous property damage resulting in enormous treatment and
repair costs. It is sometimes referred to as the "Super Termite" because:
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It has large colonies.
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The territory of a single colony can be up to 300 feet.
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It infests a wide variety of structures (including boats and
high-rise condominiums).
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It eats wood at a rapid rate.
Where Did It Come From And Where Is It Now?
The Formosan subterranean termite has been transported worldwide
from its native range in southern China to Formosa (Taiwan, where it
gets its name) and Japan. Within the last 100 years it has become
established in South Africa, Hawaii and the continental United
States. The first record of Formosan termites in the
continental United States was in 1957 from Charleston, South
Carolina. In 1965 it was found in a shipyard in Houston, Texas and
within a few years colonies were discovered in Galveston, Texas, and
New Orleans, Louisiana. It is believed that the Formosan
subterranean termite was transported to port cities on the Gulf of
Mexico and southeast Atlantic coast by ships returning from the
Pacific Theater at the end of WWII. Currently, Formosan termites are
found in Alabama, Georgia, Florida, Hawaii, Louisiana, Mississippi,
North and South Carolina, Tennessee, and Texas, while an isolated
population was found near San Diego, CA in 1992 (Fig. 1).
In Florida, they have been found in Crystal River, Dania, Ft.
Lauderdale, Hallandale, Hollywood, Jensen Beach, Jupiter, Miami,
Orlando, Palm Beach Gardens, Tampa, Tallahassee, Ft. Walton Beach,
Pensacola, and several other western panhandle communities (Fig. 2).
Within the last two years, they have been discovered in Ocala
(Marion Co.), Jacksonville, Trinity (Pasco Co.), Marco Island
(Collier Co.), Bonita Springs (Lee Co.) Debary (Volusia Co.), Cape
Coral (Lee County), and Interlacheon (Putnam Co.).
The Formosan subterranean termite has the potential to spread
throughout the whole state. Young colonies living in potted plants,
landscape timbers, or infested trees can easily be transported. A
good example is given in Louisiana and Georgia where their spread
was aided by homeowners who accidently brought termites to their
homes by buying infested railroad ties at home and garden shops.
Alates (winged reproductives) also can swarm from infested boats
into new locations as the boats travel from place to place.
The Life Of A Formosan Subterranean Termite
In Florida, Formosan termite swarms usually occur from April
through July on calm, warm, and humid evenings. Swarms are quite
large with up to tens of thousands of alates. The swarmers are
attracted to lights and are often found around windows, light
fixtures, windowsills, and spider webs in lighted areas. After
swarming and landing on the ground, the alates break off their wings
and search for a mate. Once a mate is found, the male and female
search for a crevice in damp ground or wood, hollow out a small
chamber, and crawl inside. The pair, now known as the king and
queen, mate and within a few days the queen starts laying eggs. The
young, known as larvae, hatch from the eggs and are fed by the king
and queen.
A mature colony contains distinct groups called castes (Fig. 3).
These castes look different from one another and each has a special
duty within the colony. The king and queen are the primary
reproductives and are responsible for reproduction. If the queen or
king dies or the colony becomes large, secondary reproductives may
form and begin reproduction. Soldiers defend the colony against
predators and other natural enemies. Workers take care of and feed
the larvae, reproductives and soldiers, tend the eggs, build and
maintain the nest, and search for food. Alate nymphs become alates
when they are fully grown.
Formosan subterranean termite colonies are much larger than those
of native subterranean termite species. Some have been estimated to
have over 8 million individuals compared with about 1 million
termites in large native subterranean termite colonies.
Like many other termites, the Formosan termite feeds on wood and
other materials that contain cellulose which is the main structural
component of plants. Bacteria and other single-celled organisms live
in the termite digestive system and digest cellulose providing
nutrition and energy for these termites.
Do Formosan Subterranean Termites Eat Anything Else Besides
Wood?
Although they feed mostly on wood, they will eat other
cellulose-containing materials such as cardboard and paper. However,
they are known to chew through foam insulation boards, thin lead and
copper sheeting, plaster, asphalt, and some plastics.
Is It True They Eat Concrete?
Contrary to popular myth, FORMOSAN SUBTERRANEAN TERMITES DO
NOT EAT CONCRETE nor can the soldier's defensive fluid dissolve
holes in concrete. These rumors continue because Formosan
subterranean termites are always digging through the soil. Because
of this continuous activity, they are likely to find cracks and
crevices in concrete or mortar and gain entry to a structure. This
can fool someone into thinking that Formosan subterranean termites
can eat through solid concrete.
Where Do They Live?
Subterranean termite species, such as the Formosan termite,
generally live underground. They tunnel through the soil in search
of food. Unlike native subterranean termites, Formosan termites
build large nests. These are made of carton, a hard material the
workers make from soil, chewed wood or plant matter, and their own
saliva and feces. Carton nests are quite impressive - a large,
rock-like mass constructed by hundreds of thousands or millions of
termites.
Although nesting mostly below ground, some Formosan termite
colonies will build above-ground nests that are not connected to the
soil. Nests can be made in structures where the temperature does not
get too hot or cold and there is plenty of moisture. Sources of
moisture include:
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plumbing, water heater, and roof leaks
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condensation from air conditioning units
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poor drainage from gutters and flat roofs
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seepage and rainfall on boats and ships
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porches, balconies, rooftops, etc. with plants or landscaping
that are frequently watered
What Do Formosan Subterranean Termites Look Like?
Formosan subterranean termite workers look similar to those of
native subterranean species. It is almost impossible to positively
identify Formosan termites with just the workers but the soldiers
and alates look very different and are easy to identify.
Soldier
Soldiers have orange-brown, oval-shaped heads that are quite
different from the more rectangular, straight-sided head of native
subterranean termite soldiers (Fig. 4). There is a small pore,
called the fontanelle, on the front of the head. The soldiers
produce droplets of a white, glue-like fluid from this pore when
they are attacked. This fluid gums up and disables attackers.
Soldiers have black, sickle-shaped mandibles (jaws) that can be
crossed to form an X. The bodies are yellowish-white and are about
1/5 to 1/4 inches long. Formosan subterranean termite soldiers are
very aggressive. They will even attack fingers or tools if provoked,
although their bite and fluid is harmless to humans.
Alate ("Swarmer")
Swarmers are yellowish-brown with golden brown heads, a pair of
black eyes and 2 pairs of wings of equal length. They are about 1/2
to 3/5 inch long from head to wingtip. The wings are clear with two
heavily thickened veins on the leading edge (Fig. 5) and are covered
with small hairs. These hairs are clearly visible under
magnification (Fig. 6).
How Can I Get Them Identified?
If you are not sure if the termites you have are Formosan, you
can send or bring soldiers and alates to your County Extension
office for identification. Here are some tips for sending or
bringing your samples:
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Preserve the termites in rubbing alcohol and keep them in a
small, non-breakable container with a tight-fitting, leak-proof
cap or lid.
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Termites mailed in envelopes or sandwich bags dry out, get
crushed, and break apart if they are not first put into a sturdy
container. This often makes identification difficult.
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Do not stick termites on tape or tape them to paper. This makes
identification difficult.
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Be sure to include the wings if you have alates.
Remember, the better the condition your samples are in, the
easier it will be to identify them.
How Do I Know If My House Is Infested?
Large carton nests in trees, attics, wall voids, etc., are
obvious signs of an infestation. Sometimes the damage caused by
Formosan subterranean termites is not so obvious. Soft spots, damp
or moist patches, bulges, and blistered paint or wallpaper in walls,
doors, floors, and other areas may indicate termite activity
underneath. Probing these areas with a screwdriver may reveal
damaged wood, soil, carton, and live termites. Formosan subterranean
termite infestations are recognized by the presence of lots of
soldiers.
Subterranean termites crawling above ground build mud shelter
tubes because they do not like being exposed in the open to light
and air. The tubes keep the termites from drying out and shield them
from predators, such as ants, and natural enemies. Shelter tubes are
often found on walls coming up from the ground or floor (Fig. 7).
They may also be found sticking out or dropping down to the ground
in crawl spaces, under porches or stairways, etc. Tubes and carton
may be in places where they are difficult to see such as stucco or
plaster cracks, tree holes, tree crotches, etc. You can break the
tubes open to check for termites.
Remember that Formosan subterranean termites swarm during the
evening from April through July. Keep in mind that swarmers outside
around your home could be emerging and flying in from somewhere
else. Check carefully around the premises to see if they are coming
from your property.
What Should I Do If I Have Formosan Subterranean Termites?
The best thing to do is to have the infestation professionally
treated. There are two types of control available: soil termiticides
and baits. The treatment used depends largely on the type and size
of the infestation, and which one the homeowner is most comfortable
with.
Soil Termiticides
Pre-treatment. Treating soil with a liquid termiticide
creates a chemical barrier beneath the structure (Fig. 8). Depending
on the chemical, the termites will either avoid tunneling through
treated soil or die soon after they come in contact with it. Soil
termiticides have been the standard preventive treatment for
subterranean termites up until the mid-1990s. Termiticides are
applied before the foundation slab of a structure is poured. Under
ideal conditions, protection should last from 5 to 7 years; but
under less than ideal conditions or because of improper application
it can be much less. The slightest break in the protective barrier
is all that is needed for termites to reach a structure. They can
tunnel through areas in the soil where no termiticide is present.
Expansion joints, cracks, and utility and plumbing lines are common
termite entry points through a concrete slab. Termiticide breakdown,
soil erosion, improper application, and careless construction
practices (such as leaving wooden grade stakes in the slab or
disturbing treated soil) are several ways that the chemical barrier
can be broken.
Post-construction treatment. When infestations occur after
a structure has been built, termiticides are applied by one of three
methods: rodding, drilling, or trenching. In the first, termiticide
is injected directly into the soil at specific intervals around the
perimeter of the house and beneath the slab with a rodder, an
injection tool with a long, hollow, metal rod with an open tip.
Drilling involves making holes through concrete slabs, walkways,
patios, walls, and floors in order to treat the soil beneath the
slab or inside wall voids. Trenching involves digging a shallow
trench (about 6 X 6 inches) around the base of the home, applying
termiticide to the trench and the backfill and then refilling the
trench.
Baits
Baiting systems provide an alternative to liquid termiticides.
Developed in the early 1990s, they are also effective against the
Formosan subterranean termite. Baiting involves placing bait
stations (Fig. 9) in the soil around the outside of the house. The
stations contain small pieces of wood (in some products the stations
are installed with both wood and bait) and are checked regularly for
termites. When termites are found in a station, the wood is removed
and replaced with the bait. The bait is either a paper- or
cardboard-like material or textured cellulose that contains a
substance that slowly kills the termites. The idea behind baiting is
that the termites feed on the bait and get a dose of the active
ingredient. Although this does not kill the termites immediately, it
gives them enough time to feed the other termites in the colony.
Eventually, all the members of the colony are affected. The termites
begin dying and the population of the colony is severely reduced or
eliminated.
Several different baiting systems are now being used by pest
management professionals or are commercially available. Some have
insect growth regulators (known as IGRs) as their active ingredient
(AI). These are chemical compounds that act like termite hormones
and keep the termites from developing normally. Other AIs prevent
the termites from getting energy from their food.
Aboveground bait stations are also available and are used when
termites are found in walls, doors, posts, flooring, etc. The
stations are placed directly on areas where termites are present so
that they can begin feeding immediately on the bait.
Advantages of baiting are:
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It is non-invasive (the baits are odorless and no liquid is
involved so the soil remains pesticide-free).
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Technicians usually do not need to enter the house (unless
in-ground stations are needed indoors).
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Drilling through floors and walls is usually not required.
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There is no exposure to the active ingredient because it is
self-contained within the bait station.
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Only a small amount of an active ingredient (sometimes less
than 1/20 oz.) is used for an entire treatment.
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The active ingredients are relatively harmless to humans and so
little is used it makes it even safer.
The main disadvantage is that control is not immediate. It may
take from several months to over a year to rid the home of
termites.
How Can I Keep My Home From Being Infested?
There are numerous ways you can reduce the chance of your home
being infested by Formosan and other subterranean termites:
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Remove any wood or cellulose-containing material (such as
cardboard) that is in direct contact with bare ground.
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Carefully inspect wooden items, especially railroad ties, for
termites before buying them.
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Do not leave wooden items such as planters, tubs, trellises,
railroad ties, firewood, and stakes on top of or in bare
ground.
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Anchor wooden posts for fences, decks, porches, sheds, etc. in
cement so that no wood is contacting bare ground.
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Structural wood at or near ground level should be
pressure-treated with a wood preservative. Preservatives mainly
protect against wood-decaying fungi but are also effective against
termites.
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Maintain a zone of at least one foot around the outside of your
home that is clear of plants and other landscaping materials. This
reduces soil moisture and makes it easier to inspect for shelter
tubes coming up from the ground.
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Install rain gutters to prevent water from dripping down around
the perimeter of your home.
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Keep rain gutters clear so that water drains quickly and does
not accumulate and soak the upper walls and roof of your home.
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Fix or replace leaky outdoor faucets and water lines.
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Gutter downspouts and air conditioner condensate lines should
empty out at least one foot away from the base of the home.
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The ground next to your home should slope away so that water
does not pool next to it.
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Keep sprinklers from wetting the walls of your home.
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Fix leaks in the basement, roof, water heater, appliances, and
other sources inside your home. These leaks moisten wood and
create damp environments that Formosan and other subterranean
termites like to live in.
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Remove all wooden grade stakes, form boards, supports, and
scrap wood after finishing construction or remodeling.
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Remove dead trees and plants including the roots and stumps, if
possible, from your yard.
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Eliminate or reduce the use of mulch and wood chips around the
foundation of your home. This eliminates cooler and moist soil
conditions favored by Formosan and other subterranean
termites.
Also, as part of the Florida Building Code, a standard Termite
Protection Code for new construction requires building designs and
construction practices that should reduce termite problems and make
it easier to find infestations.